Current and recent relationship evaluations offer important information about the strength and stability of relationships. The diagnostic value of relationship evaluations should motivate people to accurately recall recent changes in relationship quality (tracking accuracy) but may also increase sensitivity to times of low relationship quality (directional bias). Such a vigilant pattern of tracking accuracy and negatively biased memories of relationship quality should be amplified for those high in attachment anxiety. The current study used repeated assessments to compare memories of relationship quality across several weeks with actual evaluations of relationship quality gathered in prior weeks (787 weekly reports, N = 113, 94% women). People high in attachment anxiety more accurately tracked the ups and downs of past relationship quality, but also remembered times of low relationship quality to be lower than actually reported. People high in attachment anxiety also experienced greater depressed moods across weeks when they remembered their relationship quality more negatively. These results have important implications for understanding inconsistencies in prior research and the links between attachment anxiety and depressed mood.